Tufted wind-dispersed fagalean fruits, Pogonokarydion gen. nov., from the Upper Eocene Florissant Formation of Colorado, USA

Tufted wind-dispersed fagalean fruits, Pogonokarydion gen. nov., from the Upper Eocene Florissant Formation of Colorado, USA

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Wind-dispersed comose fruits from the Late Eocene of Florissant, Colorado, USA, are recognized as a new genus of fagalean affinity.Pogonokarydion crassulum (Cockerell) gen.et comb.nov.

disseminules consist of an ellipsoidal to circular nut bearing longitudinal ribs, a persistent epigynous perianth, paired styles, and a prominent tuft of straight hairs Tub Bearing arising from the base, which we infer to be homologous with an involucre.Prior Coffee / Espresso / Tea Machine Cleaners assignments to Juncus (Juncaceae) and Fagopsis (Fagaceae) are rejected and we argue for a position within the Fagales, likely within, or close to, the Betulaceae.Pogonokarydion co-occurred in the Florissant flora with other fagalean genera, including some still living today, like Quercus and Carya , and others now extinct like Fagopsis and Asterocarpinus.